Table of Contents

Biological Sciences

• Agricultural Sciences

  • Open Access

    Analysis of tandem gene copies in maize chromosomal regions reconstructed from long sequence reads

    • Inaugural Article

    Jiaqiang Dong, Yaping Feng, Dibyendu Kumar, Wei Zhang, Tingting Zhu, Ming-Cheng Luo, and Joachim Messing

    PNAS July 19, 2016 113 (29) 7949-7956; first published June 27, 2016. https://doi.org/10.1073/pnas.1608775113

    See companion article:

    • Profile of Joachim Messing

      - Jul 05, 2016

• Anthropology

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    Human phylogeography and diversity

    Alexander H. Harcourt

    PNAS July 19, 2016 113 (29) 8072-8078; first published July 18, 2016. https://doi.org/10.1073/pnas.1601068113

• Applied Biological Sciences

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    Dendrimer-RNA nanoparticles generate protective immunity against lethal Ebola, H1N1 influenza, and Toxoplasma gondii challenges with a single dose

    Jasdave S. Chahal, Omar F. Khan, Christopher L. Cooper, Justine S. McPartlan, Jonathan K. Tsosie, Lucas D. Tilley, Saima M. Sidik, Sebastian Lourido, Robert Langer, Sina Bavari, Hidde L. Ploegh, and Daniel G. Anderson

    PNAS July 19, 2016 113 (29) E4133-E4142; first published July 5, 2016. https://doi.org/10.1073/pnas.1600299113
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    In situ characterization of the mTORC1 during adipogenesis of human adult stem cells on chip

    Xuanye Wu, Nils Schneider, Alina Platen, Indranil Mitra, Matthias Blazek, Roland Zengerle, Roland Schüle, and Matthias Meier

    PNAS July 19, 2016 113 (29) E4143-E4150; first published July 5, 2016. https://doi.org/10.1073/pnas.1601207113
  • Open Access

    Probe for the measurement of cell surface pH in vivo and ex vivo

    Michael Anderson, Anna Moshnikova, Donald M. Engelman, Yana K. Reshetnyak, and Oleg A. Andreev

    PNAS July 19, 2016 113 (29) 8177-8181; first published July 5, 2016. https://doi.org/10.1073/pnas.1608247113

• Biochemistry

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    Structure and function of the yeast listerin (Ltn1) conserved N-terminal domain in binding to stalled 60S ribosomal subunits

    Selom K. Doamekpor, Joong-Won Lee, Nathaniel L. Hepowit, Cheng Wu, Clement Charenton, Marilyn Leonard, Mario H. Bengtson, Kanagalaghatta R. Rajashankar, Matthew S. Sachs, Christopher D. Lima, and Claudio A. P. Joazeiro

    PNAS July 19, 2016 113 (29) E4151-E4160; first published July 6, 2016. https://doi.org/10.1073/pnas.1605951113
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    Assignment of function to a domain of unknown function: DUF1537 is a new kinase family in catabolic pathways for acid sugars

    Xinshuai Zhang, Michael S. Carter, Matthew W. Vetting, Brian San Francisco, Suwen Zhao, Nawar F. Al-Obaidi, Jose O. Solbiati, Jennifer J. Thiaville, Valérie de Crécy-Lagard, Matthew P. Jacobson, Steven C. Almo, and John A. Gerlt

    PNAS July 19, 2016 113 (29) E4161-E4169; first published July 11, 2016. https://doi.org/10.1073/pnas.1605546113
  • Open Access

    Insulin induction of SREBP-1c in rodent liver requires LXRα-C/EBPβ complex

    Jing Tian, Joseph L. Goldstein, and Michael S. Brown

    PNAS July 19, 2016 113 (29) 8182-8187; first published July 5, 2016. https://doi.org/10.1073/pnas.1608987113
  • Open Access

    Resistance mutations generate divergent antibiotic susceptibility profiles against translation inhibitors

    Alexis I. Cocozaki, Roger B. Altman, Jian Huang, Ed T. Buurman, Steven L. Kazmirski, Peter Doig, D. Bryan Prince, Scott C. Blanchard, Jamie H. D. Cate, and Andrew D. Ferguson

    PNAS July 19, 2016 113 (29) 8188-8193; first published July 5, 2016. https://doi.org/10.1073/pnas.1605127113

• Biophysics and Computational Biology

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    Structural basis for sulfation-dependent self-glycan recognition by the human immune-inhibitory receptor Siglec-8

    Johannes M. Pröpster, Fan Yang, Said Rabbani, Beat Ernst, Frédéric H.-T. Allain, and Mario Schubert

    PNAS July 19, 2016 113 (29) E4170-E4179; first published June 29, 2016. https://doi.org/10.1073/pnas.1602214113
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    Generalized nucleation and looping model for epigenetic memory of histone modifications

    Fabian Erdel and Eric C. Greene

    PNAS July 19, 2016 113 (29) E4180-E4189; first published July 5, 2016. https://doi.org/10.1073/pnas.1605862113
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    Unfolding the mechanism of the AAA+ unfoldase VAT by a combined cryo-EM, solution NMR study

    Rui Huang, Zev A. Ripstein, Rafal Augustyniak, Michal Lazniewski, Krzysztof Ginalski, Lewis E. Kay, and John L. Rubinstein

    PNAS July 19, 2016 113 (29) E4190-E4199; first published July 11, 2016. https://doi.org/10.1073/pnas.1603980113
  • Open Access

    Cellular heterogeneity mediates inherent sensitivity–specificity tradeoff in cancer targeting by synthetic circuits

    Mathieu Morel, Roman Shtrahman, Varda Rotter, Lior Nissim, and Roy H. Bar-Ziv

    PNAS July 19, 2016 113 (29) 8133-8138; first published July 6, 2016. https://doi.org/10.1073/pnas.1604391113
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    Cytosine deamination and the precipitous decline of spontaneous mutation during Earth's history

    Charles A. Lewis Jr., Jesse Crayle, Shuntai Zhou, Ronald Swanstrom, and Richard Wolfenden

    PNAS July 19, 2016 113 (29) 8194-8199; first published July 5, 2016. https://doi.org/10.1073/pnas.1607580113
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    Stochastic cycle selection in active flow networks

    Francis G. Woodhouse, Aden Forrow, Joanna B. Fawcett, and Jörn Dunkel

    PNAS July 19, 2016 113 (29) 8200-8205; first published July 5, 2016. https://doi.org/10.1073/pnas.1603351113
  • Open Access

    Spectral discrimination in color blind animals via chromatic aberration and pupil shape

    Alexander L. Stubbs and Christopher W. Stubbs

    PNAS July 19, 2016 113 (29) 8206-8211; first published July 5, 2016. https://doi.org/10.1073/pnas.1524578113

    See companion article:

    • All color vision is more difficult in turbid water

      - Nov 01, 2016
  • Open Access

    Structure of human Niemann–Pick C1 protein

    • From the Cover

    Xiaochun Li, Jiawei Wang, Elias Coutavas, Hang Shi, Qi Hao, and Günter Blobel

    PNAS July 19, 2016 113 (29) 8212-8217; first published June 15, 2016. https://doi.org/10.1073/pnas.1607795113

    See companion article:

    • NPC1 structure revealed

      - Jul 07, 2016
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    Phosphotyrosine-mediated LAT assembly on membranes drives kinetic bifurcation in recruitment dynamics of the Ras activator SOS

    William Y. C. Huang, Qingrong Yan, Wan-Chen Lin, Jean K. Chung, Scott D. Hansen, Sune M. Christensen, Hsiung-Lin Tu, John Kuriyan, and Jay T. Groves

    PNAS July 19, 2016 113 (29) 8218-8223; first published July 1, 2016. https://doi.org/10.1073/pnas.1602602113
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    Dynamics and architecture of the NRBF2-containing phosphatidylinositol 3-kinase complex I of autophagy

    Lindsey N. Young, Kelvin Cho, Rosalie Lawrence, Roberto Zoncu, and James H. Hurley

    PNAS July 19, 2016 113 (29) 8224-8229; first published July 6, 2016. https://doi.org/10.1073/pnas.1603650113
  • Open Access

    Dimer interface of bovine cytochrome c oxidase is influenced by local posttranslational modifications and lipid binding

    Idlir Liko, Matteo T. Degiacomi, Shabaz Mohammed, Shinya Yoshikawa, Carla Schmidt, and Carol V. Robinson

    PNAS July 19, 2016 113 (29) 8230-8235; first published June 30, 2016. https://doi.org/10.1073/pnas.1600354113

• Cell Biology

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    DNA binding triggers tetramerization of the glucocorticoid receptor in live cells

    Diego M. Presman, Sourav Ganguly, R. Louis Schiltz, Thomas A. Johnson, Tatiana S. Karpova, and Gordon L. Hager

    PNAS July 19, 2016 113 (29) 8236-8241; first published July 5, 2016. https://doi.org/10.1073/pnas.1606774113

• Developmental Biology

  • Open Access

    Epithelial Notch signaling regulates lung alveolar morphogenesis and airway epithelial integrity

    Po-Nien Tsao, Chisa Matsuoka, Shu-Chen Wei, Atsuyasu Sato, Susumu Sato, Koichi Hasegawa, Hung-kuan Chen, Thai-Yen Ling, Munemasa Mori, Wellington V. Cardoso, and Mitsuru Morimoto

    PNAS July 19, 2016 113 (29) 8242-8247; first published June 30, 2016. https://doi.org/10.1073/pnas.1511236113

• Ecology

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    Global biogeography of microbial nitrogen-cycling traits in soil

    Michaeline B. Nelson, Adam C. Martiny, and Jennifer B. H. Martiny

    PNAS July 19, 2016 113 (29) 8033-8040; first published July 18, 2016. https://doi.org/10.1073/pnas.1601070113
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    Bison phylogeography constrains dispersal and viability of the Ice Free Corridor in western Canada

    Peter D. Heintzman, Duane Froese, John W. Ives, André E. R. Soares, Grant D. Zazula, Brandon Letts, Thomas D. Andrews, Jonathan C. Driver, Elizabeth Hall, P. Gregory Hare, Christopher N. Jass, Glen MacKay, John R. Southon, Mathias Stiller, Robin Woywitka, Marc A. Suchard, and Beth Shapiro

    PNAS July 19, 2016 113 (29) 8057-8063; first published June 6, 2016. https://doi.org/10.1073/pnas.1601077113
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    Large scale, synchronous variability of marine fish populations driven by commercial exploitation

    Kenneth T. Frank, Brian Petrie, William C. Leggett, and Daniel G. Boyce

    PNAS July 19, 2016 113 (29) 8248-8253; first published July 5, 2016. https://doi.org/10.1073/pnas.1602325113

• Evolution

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    Neighboring genes for DNA-binding proteins rescue male sterility in Drosophila hybrids

    Marjorie A. Liénard, Luciana O. Araripe, and Daniel L. Hartl

    PNAS July 19, 2016 113 (29) E4200-E4207; first published June 29, 2016. https://doi.org/10.1073/pnas.1608337113
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    Comparative phylogeography of the ocean planet

    Brian W. Bowen, Michelle R. Gaither, Joseph D. DiBattista, Matthew Iacchei, Kimberly R. Andrews, W. Stewart Grant, Robert J. Toonen, and John C. Briggs

    PNAS July 19, 2016 113 (29) 7962-7969; first published July 18, 2016. https://doi.org/10.1073/pnas.1602404113
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    Comparative phylogeography clarifies the complexity and problems of continental distribution that drove A. R. Wallace to favor islands

    Brett R. Riddle

    PNAS July 19, 2016 113 (29) 7970-7977; first published July 18, 2016. https://doi.org/10.1073/pnas.1601072113
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    Inferring responses to climate dynamics from historical demography in neotropical forest lizards

    Ivan Prates, Alexander T. Xue, Jason L. Brown, Diego F. Alvarado-Serrano, Miguel T. Rodrigues, Michael J. Hickerson, and Ana C. Carnaval

    PNAS July 19, 2016 113 (29) 7978-7985; first published July 18, 2016. https://doi.org/10.1073/pnas.1601063113
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    Comparative phylogeography of oceanic archipelagos: Hotspots for inferences of evolutionary process

    Kerry L. Shaw and Rosemary G. Gillespie

    PNAS July 19, 2016 113 (29) 7986-7993; first published July 18, 2016. https://doi.org/10.1073/pnas.1601078113
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    Effects of the population pedigree on genetic signatures of historical demographic events

    John Wakeley, Léandra King, and Peter R. Wilton

    PNAS July 19, 2016 113 (29) 7994-8001; first published July 18, 2016. https://doi.org/10.1073/pnas.1601080113
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    The probability of monophyly of a sample of gene lineages on a species tree

    Rohan S. Mehta, David Bryant, and Noah A. Rosenberg

    PNAS July 19, 2016 113 (29) 8002-8009; first published July 18, 2016. https://doi.org/10.1073/pnas.1601074113
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    Phylogeographic model selection leads to insight into the evolutionary history of four-eyed frogs

    Maria Tereza C. Thomé and Bryan C. Carstens

    PNAS July 19, 2016 113 (29) 8010-8017; first published July 18, 2016. https://doi.org/10.1073/pnas.1601064113
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    Toward a paradigm shift in comparative phylogeography driven by trait-based hypotheses

    Anna Papadopoulou and L. Lacey Knowles

    PNAS July 19, 2016 113 (29) 8018-8024; first published July 18, 2016. https://doi.org/10.1073/pnas.1601069113
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    Reticulation, divergence, and the phylogeography–phylogenetics continuum

    Scott V. Edwards, Sally Potter, C. Jonathan Schmitt, Jason G. Bragg, and Craig Moritz

    PNAS July 19, 2016 113 (29) 8025-8032; first published July 18, 2016. https://doi.org/10.1073/pnas.1601066113
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    Phenotypes in phylogeography: Species’ traits, environmental variation, and vertebrate diversification

    Kelly R. Zamudio, Rayna C. Bell, and Nicholas A. Mason

    PNAS July 19, 2016 113 (29) 8041-8048; first published July 18, 2016. https://doi.org/10.1073/pnas.1602237113
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    Geogenetic patterns in mouse lemurs (genus Microcebus) reveal the ghosts of Madagascar's forests past

    Anne D. Yoder, C. Ryan Campbell, Marina B. Blanco, Mario dos Reis, Jörg U. Ganzhorn, Steven M. Goodman, Kelsie E. Hunnicutt, Peter A. Larsen, Peter M. Kappeler, Rodin M. Rasoloarison, José M. Ralison, David L. Swofford, and David W. Weisrock

    PNAS July 19, 2016 113 (29) 8049-8056; first published July 18, 2016. https://doi.org/10.1073/pnas.1601081113
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    Evolutionary lessons from California plant phylogeography

    Victoria L. Sork, Paul F. Gugger, Jin-Ming Chen, and Silke Werth

    PNAS July 19, 2016 113 (29) 8064-8071; first published July 18, 2016. https://doi.org/10.1073/pnas.1602675113
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    Union of phylogeography and landscape genetics

    Leslie J. Rissler

    PNAS July 19, 2016 113 (29) 8079-8086; first published July 18, 2016. https://doi.org/10.1073/pnas.1601073113
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    Sympatric speciation of spiny mice, Acomys, unfolded transcriptomically at Evolution Canyon, Israel

    Kexin Li, Huihua Wang, Zhenyuan Cai, Liuyang Wang, Qinqin Xu, Matěj Lövy, Zhenlong Wang, and Eviatar Nevo

    PNAS July 19, 2016 113 (29) 8254-8259; first published July 1, 2016. https://doi.org/10.1073/pnas.1608743113
  • Open Access

    Source–sink plasmid transfer dynamics maintain gene mobility in soil bacterial communities

    James P. J. Hall, A. Jamie Wood, Ellie Harrison, and Michael A. Brockhurst

    PNAS July 19, 2016 113 (29) 8260-8265; first published July 6, 2016. https://doi.org/10.1073/pnas.1600974113
  • Open Access

    Temperate phages both mediate and drive adaptive evolution in pathogen biofilms

    Emily V. Davies, Chloe E. James, David Williams, Siobhan O’Brien, Joanne L. Fothergill, Sam Haldenby, Steve Paterson, Craig Winstanley, and Michael A. Brockhurst

    PNAS July 19, 2016 113 (29) 8266-8271; first published July 5, 2016. https://doi.org/10.1073/pnas.1520056113

• Immunology and Inflammation

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    pH-degradable imidazoquinoline-ligated nanogels for lymph node-focused immune activation

    Lutz Nuhn, Nane Vanparijs, Ans De Beuckelaer, Lien Lybaert, Glenn Verstraete, Kim Deswarte, Stefan Lienenklaus, Nikunj M. Shukla, Alex C. D. Salyer, Bart N. Lambrecht, Johan Grooten, Sunil A. David, Stefaan De Koker, and Bruno G. De Geest

    PNAS July 19, 2016 113 (29) 8098-8103; first published July 5, 2016. https://doi.org/10.1073/pnas.1600816113
  • Open Access

    Identification of shared TCR sequences from T cells in human breast cancer using emulsion RT-PCR

    • From the Cover

    Daniel J. Munson, Colt A. Egelston, Kami E. Chiotti, Zuly E. Parra, Tullia C. Bruno, Brandon L. Moore, Taizo A. Nakano, Diana L. Simons, Grecia Jimenez, John H. Yim, Dmitri V. Rozanov, Michael T. Falta, Andrew P. Fontenot, Paul R. Reynolds, Sonia M. Leach, Virginia F. Borges, John W. Kappler, Paul T. Spellman, Peter P. Lee, and Jill E. Slansky

    PNAS July 19, 2016 113 (29) 8272-8277; first published June 15, 2016. https://doi.org/10.1073/pnas.1606994113

    See companion article:

    • Shared antigens in cancer immunotherapy

      - Jul 07, 2016
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    CCR7 expression alters memory CD8 T-cell homeostasis by regulating occupancy in IL-7– and IL-15–dependent niches

    Yong Woo Jung, Hyun Gyung Kim, Curtis J. Perry, and Susan M. Kaech

    PNAS July 19, 2016 113 (29) 8278-8283; first published July 6, 2016. https://doi.org/10.1073/pnas.1602899113
  • Open Access

    Vagus nerve stimulation inhibits cytokine production and attenuates disease severity in rheumatoid arthritis

    Frieda A. Koopman, Sangeeta S. Chavan, Sanda Miljko, Simeon Grazio, Sekib Sokolovic, P. Richard Schuurman, Ashesh D. Mehta, Yaakov A. Levine, Michael Faltys, Ralph Zitnik, Kevin J. Tracey, and Paul P. Tak

    PNAS July 19, 2016 113 (29) 8284-8289; first published July 5, 2016. https://doi.org/10.1073/pnas.1605635113

• Medical Sciences

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    Ascorbate attenuates pulmonary emphysema by inhibiting tobacco smoke and Rtp801-triggered lung protein modification and proteolysis

    Indranil Gupta, Souradipta Ganguly, Christine R. Rozanas, Dennis J. Stuehr, and Koustubh Panda

    PNAS July 19, 2016 113 (29) E4208-E4217; first published July 5, 2016. https://doi.org/10.1073/pnas.1600056113
  • Open Access

    Sleeping Beauty screen reveals Pparg activation in metastatic prostate cancer

    Imran Ahmad, Ernest Mui, Laura Galbraith, Rachana Patel, Ee Hong Tan, Mark Salji, Alistair G. Rust, Peter Repiscak, Ann Hedley, Elke Markert, Carolyn Loveridge, Louise van der Weyden, Joanne Edwards, Owen J. Sansom, David J. Adams, and Hing Y. Leung

    PNAS July 19, 2016 113 (29) 8290-8295; first published June 29, 2016. https://doi.org/10.1073/pnas.1601571113

• Microbiology

  • Open Access

    piRNA pathway is not required for antiviral defense in Drosophila melanogaster

    Marine Petit, Vanesa Mongelli, Lionel Frangeul, Hervé Blanc, Francis Jiggins, and Maria-Carla Saleh

    PNAS July 19, 2016 113 (29) E4218-E4227; first published June 29, 2016. https://doi.org/10.1073/pnas.1607952113
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    Deletion of a dehydratase important for intracellular growth and cording renders rough Mycobacterium abscessus avirulent

    Iman Halloum, Séverine Carrère-Kremer, Mickael Blaise, Albertus Viljoen, Audrey Bernut, Vincent Le Moigne, Catherine Vilchèze, Yann Guérardel, Georges Lutfalla, Jean-Louis Herrmann, William R. Jacobs Jr., and Laurent Kremer

    PNAS July 19, 2016 113 (29) E4228-E4237; first published July 6, 2016. https://doi.org/10.1073/pnas.1605477113
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    Interbacterial signaling via Burkholderia contact-dependent growth inhibition system proteins

    • From the Cover

    Erin C. Garcia, Andrew I. Perault, Sara A. Marlatt, and Peggy A. Cotter

    PNAS July 19, 2016 113 (29) 8296-8301; first published June 22, 2016. https://doi.org/10.1073/pnas.1606323113

    See companion article:

    • Interactions between bacteria and their responses

      - Jul 15, 2016
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    Temporal and intrinsic factors of rifampicin tolerance in mycobacteria

    Kirill Richardson, Owen T. Bennion, Shumin Tan, Anh N. Hoang, Murat Cokol, and Bree B. Aldridge

    PNAS July 19, 2016 113 (29) 8302-8307; first published June 29, 2016. https://doi.org/10.1073/pnas.1600372113

• Neuroscience

  • Open Access

    Decoding and disrupting left midfusiform gyrus activity during word reading

    Elizabeth A. Hirshorn, Yuanning Li, Michael J. Ward, R. Mark Richardson, Julie A. Fiez, and Avniel Singh Ghuman

    PNAS July 19, 2016 113 (29) 8162-8167; first published June 20, 2016. https://doi.org/10.1073/pnas.1604126113

    See companion article:

    • Intracranial dissection of word reading mechanisms

      - Jul 07, 2016
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    Early multisensory integration of self and source motion in the auditory system

    Eyal Wigderson, Israel Nelken, and Yosef Yarom

    PNAS July 19, 2016 113 (29) 8308-8313; first published June 29, 2016. https://doi.org/10.1073/pnas.1522615113
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    Inositol pyrophosphates inhibit synaptotagmin-dependent exocytosis

    Tae-Sun Lee, Joo-Young Lee, Jae Won Kyung, Yoosoo Yang, Seung Ju Park, Seulgi Lee, Igor Pavlovic, Byoungjae Kong, Yong Seok Jho, Henning J. Jessen, Dae-Hyuk Kweon, Yeon-Kyun Shin, Sung Hyun Kim, Tae-Young Yoon, and Seyun Kim

    PNAS July 19, 2016 113 (29) 8314-8319; first published June 30, 2016. https://doi.org/10.1073/pnas.1521600113

• Plant Biology

  • Open Access

    Mapping transcription factor interactome networks using HaloTag protein arrays

    Junshi Yazaki, Mary Galli, Alice Y. Kim, Kazumasa Nito, Fernando Aleman, Katherine N. Chang, Anne-Ruxandra Carvunis, Rosa Quan, Hien Nguyen, Liang Song, José M. Alvarez, Shao-shan Carol Huang, Huaming Chen, Niroshan Ramachandran, Stefan Altmann, Rodrigo A. Gutiérrez, David E. Hill, Julian I. Schroeder, Joanne Chory, Joshua LaBaer, Marc Vidal, Pascal Braun, and Joseph R. Ecker

    PNAS July 19, 2016 113 (29) E4238-E4247; first published June 29, 2016. https://doi.org/10.1073/pnas.1603229113
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    Methylation interactions in Arabidopsis hybrids require RNA-directed DNA methylation and are influenced by genetic variation

    Qingzhu Zhang, Dong Wang, Zhaobo Lang, Li He, Lan Yang, Liang Zeng, Yanqiang Li, Cheng Zhao, Huan Huang, Heng Zhang, Huiming Zhang, and Jian-Kang Zhu

    PNAS July 19, 2016 113 (29) E4248-E4256; first published July 5, 2016. https://doi.org/10.1073/pnas.1607851113
  • Open Access

    Borrowed alleles and convergence in serpentine adaptation

    Brian J. Arnold, Brett Lahner, Jeffrey M. DaCosta, Caroline M. Weisman, Jesse D. Hollister, David E. Salt, Kirsten Bomblies, and Levi Yant

    PNAS July 19, 2016 113 (29) 8320-8325; first published June 29, 2016. https://doi.org/10.1073/pnas.1600405113
  • Open Access

    Grasses use an alternatively wired bHLH transcription factor network to establish stomatal identity

    Michael T. Raissig, Emily Abrash, Akhila Bettadapur, John P. Vogel, and Dominique C. Bergmann

    PNAS July 19, 2016 113 (29) 8326-8331; first published July 5, 2016. https://doi.org/10.1073/pnas.1606728113

• Psychological and Cognitive Sciences

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    Learning rational temporal eye movement strategies

    David Hoppe and Constantin A. Rothkopf

    PNAS July 19, 2016 113 (29) 8332-8337; first published July 5, 2016. https://doi.org/10.1073/pnas.1601305113