Thirty thousand-year-old evidence of plant food processing
- aIstituto Italiano di Preistoria e Protostoria, Florence 50121, Italy;
- bSoprintendenza per i Beni Archeologici per la Toscana, Florence 50121, Italy;
- cDipartimento di Scienze Ambientali “G. Sarfatti”–Unità di Ricerca Ecologia Preistorica, University of Siena and Museo Civico di Storia Naturale, Verona 37129, Italy;
- dScienze e Tecnologie Alimentari, Facoltà di Agraria, University of Molise, Società Italiana di Scienze e Tecnologie Alimentari, Campobasso 86100, Italy;
- eDipartimento di Biologia Evoluzionistica-Biologia Vegetale, University of Florence, Florence 50121, Italy;
- fInstitute for the History of Material Culture, Russian Academy of Sciences, Saint Petersburg 191186, Russia;
- gInstitute of Archaeology, Russian Academy of Sciences, Moscow 140127, Russia;
- hDepartment of Anthropology, Faculty of Science, Masaryk University, Kotlarska 2, Brno, Czech Republic; and
- iInstitute of Archaeology, Academy of Science of Czech Republic, Kralovopolska 147, Brno, Czech Republic
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Edited* by Erik Trinkaus, Washington University, St. Louis, MO, and approved September 7, 2010 (received for review May 21, 2010)

Abstract
European Paleolithic subsistence is assumed to have been largely based on animal protein and fat, whereas evidence for plant consumption is rare. We present evidence of starch grains from various wild plants on the surfaces of grinding tools at the sites of Bilancino II (Italy), Kostenki 16–Uglyanka (Russia), and Pavlov VI (Czech Republic). The samples originate from a variety of geographical and environmental contexts, ranging from northeastern Europe to the central Mediterranean, and dated to the Mid-Upper Paleolithic (Gravettian and Gorodtsovian). The three sites suggest that vegetal food processing, and possibly the production of flour, was a common practice, widespread across Europe from at least ~30,000 y ago. It is likely that high energy content plant foods were available and were used as components of the food economy of these mobile hunter–gatherers.
Footnotes
- 1To whom correspondence should be addressed. E-mail: annarevedin{at}iipp.it.
Author contributions: A.R., B.A., L.L., and M.M.L. designed research; A.R., B.A., L.L., M.M.L., A.S., and J.S. performed research; R.B., L.L., E.M., N.S., and E.S. analyzed data; and A.R., B.A., L.L., M.M.L., and J.S. wrote the paper.
The authors declare no conflict of interest.
↵*This Direct Submission article had a prearranged editor.
This article contains supporting information online at www.pnas.org/lookup/suppl/doi:10.1073/pnas.1006993107/-/DCSupplemental.