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PNAS Terms of Access: “Proceedings of the National Academy of Sciences” and “PNAS” are registered trademarks of the National Academy of Sciences of the United States of America (NAS). PNAS and PNAS Online are both copyrighted © 2018 by the National Academy of Sciences.

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This publication is an online edition of the printed edition of PNAS. This online edition may not contain all of the information and material contained in the printed edition, the information it contains may not necessarily be as current as the printed edition, and in certain instances may be inaccurate or incorrect. The online edition of PNAS officially published by the NAS is the version of record.

In no event shall the NAS be liable for any special, incidental, indirect or consequential damages of any kind, or any damages whatsoever resulting from loss of use, data or profits, whether or not advised of the possibility of damage, and on any theory of liability, arising out of or in connection with the use or performance of this information.

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